Corresponding author: Sebastian Świerszcz ( seb.swierszcz@gmail.com ) Academic editor: Wolfgang Willner
© 2021 Arkadiusz Nowak, Sebastian Świerszcz, Sylwia Nowak, Marcin Nobis.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Nowak A, Świerszcz S, Nowak S, Nobis M (2021) New syntaxa of tall-forb vegetation in the Pamir-Alai and western Tian Shan Mts. (Tajikistan and Kyrgyzstan, Middle Asia) – an addendum to Nowak et al. 2020. Vegetation Classification and Survey 2: 191-194. https://doi.org/10.3897/VCS/2021/73498
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We validate eleven syntaxa (eight associations and three alliances) of tall-forb vegetation that were published earlier as nomina provisoria according to the International Code of Phytosociological Nomenclature. The validation concerns syntaxa of tall-forb vegetation of the class Prangetea ulopterae Klein 1987 reported from Pamir-Alai and western Tian Shan Mountains (Tajikistan and Kyrgyzstan).
Taxonomic reference:
Abbreviations: ICPN = International Code of Phytosociological Nomenclature.
International Code of Phytosociological Nomenclature (ICPN), Middle Asia, phytosociological nomenclature, Prangetea ulopterae, syntaxonomy, tall forb, validation
According to the older version of the International Code of Phytosociological Nomenclature (ICPN,
As a result of the phytosociological survey in alpine and subalpine belts of Pamir Alai and Tian Shan Mts. in Middle Asia, eleven new syntaxa of tall-forb vegetation were described with the provisional names (
The aim of this paper is to validate the provisional names of tall-forb syntaxa according to the current ICPN and to present the overview of the vegetation types that were described in the first paper devoted to tall-forbs of Middle Asia (
The numbers of syntaxa refer to the synopsis at the end of the paper. Diagnostic taxa are provided in
Holotypus: Eremuretum kaufmannii Nowak et al. ass. nov. hoc loco (see below)
2.1.3. Heracleetum lehmannianii Nowak et al. ass. nov. hoc loco
Holotypus: 02 July 2017; 40.24556°N, 73.34028°E; 2157 m a.s.l.; aspect N; slope 3°; plot area 10 m2; cover herb layer 90%; cover moss layer 3%; species richness: 28.
Herb layer: Geranium regelii 3, Euphorbia lamprocarpa 3, Achillea millefolium 2, Carum carvi 2, Dactylis glomerata 2, Poa pratensis 2, Ranunculus brevirostris 2, Heracleum lehmannianum 2, Amoria repens 1, Convolvulus arvensis 1, Cousinia pseudarctium 1, Festuca rubra 1, Galium turkestanicum 1, Glycyrrhiza glabra 1, Hordeum turkestanicum 1, Ligularia thomsonii 1, Mentha asiatica 1, Nepeta cataria 1, Plantago lanceolata 1, Codonopsis clematidea +, Cynoglossum viridiflorum +, Erigeron allochrous +, Lithospermum officinale +, Polygonum coriarium +, Potentilla pedata +, Trifolium pratense +, Vicia tenuifolia +. Moss layer: Ceratodon purpureus 1. [relevé number in
2.1.4. Eremuretum kaufmannii Nowak et al. ass. nov. hoc loco
Holotypus: 23 June 2014; 39.25722°N, 68.17833°E; 2836 m a.s.l.; aspect N; slope 25°; plot area 10 m2; cover herb layer 95%; species richness: 17; species composition:
Herb layer: Geranium regelii 3, Myosotis alpestris 3, Ligularia thomsonii 2, Poa urssulensis 2, Artemisia dracunculus 1, Astragalus saratagius 1, Carex turkestanica 1, Eremurus kaufmannii 1, Pedicularis grigorjevii 1, Tulipa ingens 1, Veronica rubrifolia 1, Alopecurus pratensis +, Astragalus nuciferus +, Myosotis micrantha +, Nepeta podostachys +, Oxytropis capusii +, Ziziphora pamiroalaica +. [relevé number in
2.1.5. Anthriscidetum glacialis Nowak et al. ass. nov. hoc loco
Holotypus: 16 June 2019; 38.65522°N, 70.73306°E; 2762 m a.s.l.; aspect W; slope 10°; plot area 10 m2; cover herb layer 100%; species richness: 33.
Herb layer: Anthriscus glacialis 3, Geranium regelii 2, Ligularia thomsonii 2, Paeonia intermedia 2, Polygonum hissaricum 2, Astragalus aksuensis 1, Carex dimorphotheca 1, Crepis darvazica 1, Dactylis glomerata 1, Euphorbia glomerulans 1, Fritillaria regelii 1, Leonurus turkestanicus 1, Myosotis alpestris 1, Phleum pratense 1, Poa pratensis 1, Polygonum coriarium 1, Amoria repens +, Astragalus adpressipilosus +, Astragalus tecti-mundi +, Calamagrostis pseudophragmites +, Crocus korolkowii +, Euphorbia sarawschanica +, Galium aparine +, Ligularia alpigena +, Lolium cuneatum +, Medicago romanica +, Oberna wallichiana +, Pedicularis olgae +, Picris nuristanica +, Potentilla sericea +, Rumex nepalensis +, Scilla puschkinioides +, Taraxacum sp. +. [relevé number in
Holotypus: Eremuretum stenophyllido-comosi Nowak et al. ass. nov. hoc loco (see below)
2.2.2. Phlomoidetum kaufmannianae Nowak et al. ass. nov. hoc loco
Holotypus: 29 May 2015; 39.20556°N, 67.81694°E; 1715 m a.s.l.; aspect W; slope 15°; plot area 10 m2; cover herb layer 80%; cover moss layer 10%; species richness: 25.
Herb layer: Phlomoides kaufmanniana 3, Papaver pavoninum 2, Anisantha tectorum 1, Artemisia rutifolia 1, Crambe kotschyana 1, Crepis pulchra 1, Ferula kokanica 1, Ferula ovina 1, Piptatherum kokanicum 1, Scandix stellata 1, Taeniatherum crinitum 1, Alcea nudiflora +, Alyssum calycinum +, Arabis recta +, Arenaria serpyllifolia +, Cerastium inflatum +, Erodium cicutarium +, Galium aparine +, Galium spurium +, Geranium divaricatum +, Lappula consanguinea +, Perovskia virgata +. Moss layer: Bryum caespiticium 1, Pohlia nutans 1, Encalypta vulgaris +. [relevé number in
2.2.5. Eremuretum stenophyllido-comosi Nowak et al. ass. nov. hoc loco
Holotypus: 22 May 2019; 38.70359°N, 70.46913°E; 1667 m a.s.l.; aspect SW; slope 40°; plot area 10 m2; cover shrub layer 15%; cover herb layer 70%; species richness 40.
Shrub layer: Cerasus verrucosa 2, Rosa popovii 2. Herb layer: Eremurus comosus 2, Ferula kuhistanica 2, Prangos pabularia 2, Phlomoides lehmanniana 1, Vinca erecta +, Achillea biebersteinii r, Alcea nudiflora r, Anisantha sterilis r, Antonina debilis r, Asparagus neglectus r, Asperula setosa r, Astragalus macronyx r, Bromus lanceolatus r, Bunium persicum r, Callipeltis cucullaris r, Centaurea squarrosa r, Convolvulus arvensis r, Dianthus darvazicus r, Diarthron vesiculosum r, Dichasianthus subtilissimus r, Eremurus stenophyllus r, Euphorbia franchetii r, Euphorbia sarawschanica r, Galium spurium r, Hordeum bulbosum r, Lactuca tatarica r, Lallemantia royleana r, Lappula consanguinea r, Papaver litwinowii r, Poterium polygamum r, Rheum maximowiczii r, Rhinopetalum bucharicum r, Salvia sclarea r, Serratula chartacea r, Strigosella trichocarpa r, Tanacetum pseudachillea r, Taraxacum nuratavicum r, Veronica rubrifolia r. [relevé number in
Dry tall-forb communities of the subhumid zone of the eastern Irano-Turanian region
Holotypus: Phlomoido lehmannianae-Onobrychidetum grandis Nowak et al. ass. nov. hoc loco (see below)
2.3.3. Lathyretum mulkaki Nowak et al. ass. nov. hoc loco
Holotypus: 17 June 2019; 38.58247°N, 70.76009°E; 2640 m a.s.l.; aspect SW; slope 25°; plot area 10 m2; cover shrub layer 2%; cover herb layer 75%; species richness: 31.
Shrub layer: Rosa ovczinnikovii 1. Herb layer: Ferula kokanica 3, Dictamnus angustifolius 2, Lathyrus mulkak 2, Senecio franchetii 2, Bunium persicum 1, Centaurea ruthenica 1, Delphinium batalinii 1, Nepeta ucranica 1, Poa bulbosa 1, Scabiosa songarica 1, Thalictrum kuhistanicum 1, Ziziphora pamiroalaica 1, Allium hissaricum +, Asperula setosa +, Colchicum luteum +, Cousinia pseudarctium +, Dianthus darvazicus +, Galium spurium +, Hypericum scabrum +, Iris darwasica +, Lappula occultata +, Lophanthus elegans +, Nepeta podostachys +, Origanum tyttanthum +, Phlomoides arctifolia +, Poa zaprjagajevii +, Pseudoclausia turkestanica +, Rhinopetalum bucharicum +, Thymus seravschanicus +, Vinca erecta +. [relevé number in
2.3.7. Eremuretum robusti Nowak et al. ass. nov. hoc loco
Holotypus: 05 June 2015; 38.82000°N, 70.20778°E; 1436 m a.s.l.; plot area 10 m2; cover herb layer 100%; species richness: 23.
Herb layer: Phlomoides arctifolia 3, Vicia tenuifolia 3, Galium pamiroalaicum 2, Phlomoides lehmanniana 2, Eremurus comosus 1, Eremurus robustus 1, Eremurus stenophyllus 1, Hordeum bulbosum 1, Pimpinella peregrina 1, Poa bulbosa 1, Poa pratensis 1, Potentilla transcaspia 1, Poterium lasiocarpum 1, Prangos pabularia 1, Arenaria serpyllifolia +, Arum korolkowii +, Cousinia pseudarctium +, Galium aparine +, Orobanche sulphurea +, Plantago lanceolata +, Salvia sclarea +, Taraxacum sp. +, Torilis arvensis +. [relevé number in
2.3.8. Phlomoido lehmannianae-Onobrychidetum grandis Nowak et al. ass. nov. hoc loco
Holotypus: 14 June 2019; 38.84710°N, 71.30857°E; 2519 m a.s.l.; aspect S; slope 30°; plot area 10 m2; cover shrub layer 5%; cover herb layer 100%; species richness: 26.
Shrub layer: Rosa ovczinnikovii 2. Herb layer: Ferula kuhistanica 3, Phlomoides lehmanniana 3, Prangos pabularia 3, Onobrychis grandis 2, Scabiosa songarica 2, Carex turkestanica +, Poa bulbosa +, Alyssum calycinum r, Arenaria serpyllifolia r, Asperula setosa r, Drepanocaryum sewerzowii r, Elaeosticta allioides r, Gentiana olivieri r, Handelia trichophylla r, Hypericum scabrum r, Lappula patula r, Neurotropis kotschyana r, Poa zaprjagajevii r, Polygonum paronychioides r, Polygonum polycnemoides r, Rochelia cardiosepala r, Serratula chartacea r, Stellaria alsinoides r, Taraxacum sp. r, Veronica cardiocarpa r. [relevé number in
This overview follows the classification scheme of
Class: Molinio-Arrhenatheretea Tx. 1937
1. Order: Poo alpinae-Trisetetalia Ellmauer et Mucina 1993
1.1. Alliance: Poion alpinae Gams ex Oberd. 1950
1.1.1. Community of Phlomoides oreophila
Class: Prangetea ulopterae Klein 1987.
2.1. Alliance: Ligulario thomsonii-Geranion regelii Nowak et al. 2021 all. nov.
2.1.1. Community of Allium hymenorhizum
2.1.2. Feruletum sumbuli Nowak et al. 2015
2.1.3. Heracleetum lehmannianii Nowak et al. 2021 ass. nov.
2.1.4. Eremuretum kaufmannii Nowak et al. ass. 2021 ass. nov.
2.1.5. Anthriscidetum glacialis Nowak et al. 2021 ass. nov.
2.2. Alliance: Rheion maximoviczii Nowak et al. all. 2021 all. nov.
2.2.1. Community of Cousinia batalinii and Euphorbia pamirica
2.2.2. Phlomoidetum kaufmannianae Nowak et al. 2021 ass. nov.
2.2.3. Phlomoidetum tadshikistanicae
2.2.4. Community of Senecio saposhnikovii
2.2.5. Eremuretum stenophyllido-comosi Nowak et al. 2021 ass. nov.
2.3. Alliance: Scabioso songaricae-Phlomoidion lehmannianae Nowak et al. 2021 all. nov.
2.3.1. Community of Inula macrophylla
2.3.2. Stipetum margelanicae
2.3.3. Lathyretum mulkaki Nowak et al. 2021 ass. nov.
2.3.4. Potentillo orientalis-Eremuretum fusci Świerszcz et al. 2020
2.3.5. Hordeo bulbosi-Astragaletum retamocarpi Świerszcz et al. 2020
2.3.6. Community of Ferula kuhistanica
2.3.7. Eremuretum robusti Nowak et al. 2021 ass. nov.
2.3.8. Phlomoido lehmannianae-Onobrychidetum grandis Nowak et al. 2021 ass. nov.
A.N. led the writing, while all authors critically revised the final manuscript.
The authors are grateful to Wolfgang Willner for providing valuable comments on the text.